Business travelers paid more for flights during the second quarter this year than at any time since 2001, says a new study to be released Tuesday by American Express Business Travel.

AmEx's quarterly Business Travel Monitor showed the average airfare paid by business travelers in the April-June quarter this year jumped 13% to $247 one way from $218 in the same period in 2005.

Travelers also paid more for hotels and rental cars.

"More travelers in the air, combined with higher fuel prices and fewer available seats and hotel rooms, have led to considerable price increases - a trend we expect to continue," said Frank Schnur, vice president of consulting for American Express Business Travel.

For the moment, fares don't appear to be headed higher. But they haven't declined as much since Labor Day this year as in previous years, either.

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